Publications

Peer-Reviewed Publications

• Y. Zhao, K. Zhu, Y. Zhang, & D.B. Cooney (2026). Pattern Formation in a Spatial Public Goods Dilemma due to Diffusive or Directed Motion. Bulletin of Mathematical Biology, 88(173).
• S.M. Peyres, D.B. Cooney, D.B. Go, R.M. Sankaran, & N.B. Üner (2026). Operating Regimes, Scaling Laws, and Optimization of Selectivity in Plasma-Liquid Reactions. Chemical Engineering Science, 335, 124342.
• T. Yao*, C. Xu*, & D.B. Cooney (2026). Pattern Formation in Agent-Based and PDE Models for Evolutionary Games with Payoff-Driven Motion. Bulletin of Mathematical Biology, 88(46). (* denotes joint first authors)
• K. Alexiou & D.B. Cooney (2025). Steady-State and Dynamical Behavior of a PDE Model of Multilevel Selection with Pairwise Group-Level Competition. Bulletin of Mathematical Biology, 87(97).
• D.B. Cooney (2025). Exploring the Evolution of Altruistic Punishment with a PDE Model of Cultural Multilevel Selection. Bulletin of Mathematical Biology, 87(51), 1-99.
• D.B. Cooney, S.A. Levin, Y. Mori, and J.B. Plotkin (2023). Evolutionary Dynamics Within and Among Competing Groups. PNAS, 120(20), e2216186120.
• J. Benson, M. Bessonov, K. Burke, S. Cassani, M.V. Ciocanel, D.B. Cooney, & A. Volkening (2023). How do classroom-turnover times depend on lecture-hall size?. Mathematical Biosciences and Engineering, 20(5): 9179-9207.
• D.B. Cooney*, D.H. Morris*, S.A. Levin, D.I. Rubenstein, & P. Romanczuk (2022). Social Dilemmas of Sociality due to Beneficial and Costly Contagion. PLoS Computational Biology, 18(11): e1010670. (* joint first authors)
• D.B. Cooney (2022). Assortment and Reciprocity Mechanisms for Promotion of Cooperation in a Model of Multilevel Selection. Bulletin of Mathematical Biology, 84(11), 1-49.
• D.B. Cooney, F.W. Rossine, D.H. Morris, & S.A. Levin (2022). A PDE Model for Protocell Evolution and the Origin of Chromosomes via Multilevel Selection. Bulletin of Mathematical Biology, 84(109), 1-90.
• D.B. Cooney & Y. Mori (2022). Long-Time Behavior of a PDE Replicator Equation for Multilevel Selection in Group-Structured Populations. Journal of Mathematical Biology, 85(2), 1-67.
• R. Carmona, D.B. Cooney, C.V. Graves, & M. Laurière (2022). Stochastic Graphon Games: I. The Static Case. Mathematics of Operations Research, 47(1), 750-778.
• D.B. Cooney (2020). Analysis of Multilevel Replicator Dynamics for General Two-Strategy Social Dilemmas. Bulletin of Mathematical Biology, 82(6), 1-72.
• D.B. Cooney (2019). The Replicator Dynamics for Multilevel Selection in Evolutionary Games. Journal of Mathematical Biology, 79(1), 101-154.
• D.B. Cooney, B. Allen, & C. Veller (2016). Assortment and the Evolution of Cooperation in a Moran Process with Exponential Fitness. Journal of Theoretical Biology, 409, 38-46.
• V. Wong, D.B. Cooney, & Y. Bar-Yam (2016). Beyond Contact Tracing: Community-Based Early Detection for Ebola Response. PLoS Currents: Outbreaks.

Preprint

• D. Kim*, Y. Zhang*, Y. Zhao*, & D.B. Cooney (2026). The Keller-Segel Model for Chemotaxis on Metapopulation Graphs and Graphons. arXiv:2610.31049. (* joint first authors)
• O. Clifton, S. Dodson, & D.B. Cooney (2026). Kernel-Dependent Pattern Formation in a Population Model with Nonlocal Facilitation and Competition. arXiv:2608.23964.
• T. Yao & D.B. Cooney (2025). Spatial Pattern Formation in Eco-Evolutionary Games with Environment-Driven Motion. arXiv:2502.06723.

Expository Articles

• D.B. Cooney, S.A. Levin, Y. Mori, J.B. Plotkin. Modeling Natural Selection at Multiple Levels of Organization. SIAM News Blog, October 2021.
• D.B. Cooney. Report on SIAM DS21. DS Web: The Dynamical Systems Web, July 2021.